Brooklyn Entomological Society

VoL XLVI 1951









No. 1




Brooklyn Entomological Society




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VoL. XLVI February, 1951 No. 1


By H. J. Rein HARD, College Station, Texas.

The following descriptions of new species are based mainly upon material collected by Messrs. F. A. Cowan and M. R. Wheeler in Mexico and western United States. I am indebted to the latter for the privilege of studying their extensive collections of Muscoid flies and to Dr. George F. Knowlton, who submitted one of the forms treated herein as indicated under the description of same. Types of the new species are in my collection.

Belvosia auratilis n. sp.

A large robust species closely allied to borealis Aldrich, but the male genital forceps are distinctly longer, the palpi black and the first two abdominal segments each with only one pair of median marginals.

Male: Front at vertex 0.40 of head width, gradually widening downward into facial angle ; f rentals in three irregular rows, median one sharply divergent in three or four bristles beneath antennal base ; ocellars and orbitals absent ; verticals two pairs, stout ; para- frontal with gray pollen below becoming thinner and blackish to- wards vertex ; median vitta broad, concolorous with parafrontal ; face, parafacial, cheek and posterior orbit silvery white; third an- tennal segment black, two to two and one-half times length of sec- ond ; latter usually reddish on front margin and on apex ; arista shorter than antenna, brownish black ; thickened and tapering to tip ; vibrissae over half the length of second antennal segment above the oral margin ; parafacial broad, about twice clypeal width ; facial ridges with moderately strong bristles extending above the middle ; cheek and upper half of parafacial black-haired ; palpus stout, occa-

^ Contribution No. 1222, Department of Entomology, Texas Ag- ricultural Experiment Station.


2 Bulletin of the Brooklyn Entomological Society ^LVI

sionally paler or reddish on extreme tip ; back of head flat, cinereous, clothed with pale hairs.

Thorax subshining black, lightly dusted with grayish pollen above, vittae indistinct ; sternopleurals usually four ; scutellum with five pairs of laterals and numerous irregularly spaced discals on apical half. Legs black, stout, bristly ; front pulvillus exceeding length of last tarsal segment. Wing and calypter black ; costal spine vestigial ; apical cell open well before wing tip ; subepaulet blackish, tinged with red on inner apical margin ; epaulet wholly black.

Abdomen subshining black, third and fourth segments golden pollinose on basal three-fourths or more, each with a complete marginal row of stout bristles ; genitalia black ; forceps subequal the length of first segment of hind tarsus, rather straight in profile with tips gently bowed forward, hind surface transversely convex and moderately clothed with long black hairs ; accessory process as long as forceps and nearly as thick as latter in profile, basal half bearing numerous long, coarse hairs directed outwardly, tip broadly rounded; fifth sternite black, with a broad V-shaped median ex- cision, inner margin of lobes beset with black hairs and bristles near base.

Female: Front at vertex 0.44 of head width; two to four pro- clinate and one or two reclinate orbitals ; second antennal segment one-half length of third and usually more extensively reddish in ground color than in male.

Length, 17-18 mm.

Holotype male and allotype female, Guadalajara, Mich., Mexico, August 27, 1947 (F. A. Cowan and M. R. Wheeler). Paratypes : 3 males and 7 females, same data as type.

Belvosia ansata n. sp.

Dif¥ers from the preceding species chiefly as follows : Male smaller and less robust in build ; front at vertex 0.40 of head width ; palpus wholly red ; vibrissae situated the length of second antennal segment above oral margin ; facial ridges with stout bristles ascend- ing about to level with arista ; parafacial but slightly narrowed below and nearly equal to clypeal width on lower extremity, black hairs beneath frontals small and averaging less than six in number. Thorax black with thin grayish pollen anteriorly; scutellum deep reddish brown, normally with four pairs of lateral bristles and usually a short spinelike apical or preapical pair. Front claws and pulvilli elongated, almost equal to last two tarsal segments. Wing and calypter deep brownish to black ; subepaulet wholly orange red. Abdominal segments three and four with golden pollen extending

Fel)., 1951 Bulletin of the Brooklyn Entomological Society


to hind margin on venter but less distinctly so on sides, the upper posterior edge of each appearing blackish in direct view ; second genital segment reddish on sides ; forceps about half as long as first segment on hind tarsus, triangular in rear view ; accessory process widest at middle, tapering apically to a narrow rounded tip, sparsely clothed with fine hairs basally on outer side.

Female: Front at vertex 0.40 of head width; one reclinate and usually three proclinate orbitals; second antennal segment slightly over one-half length of third, more or less reddish in ground color ; front pulvillus a little shorter than apical tarsal segment.

Length, 14-15 mm.

Holotype male and allotype female, Guadalajara, Mich., Mexico, August 27, 1947 (F. A. Cowan and M. R. Wheeler). Paratypes: 1 1 males and 4 females, same data as type.

The species resembles bifasciata in general appearance but is readily distinguished in having the calypter black, subepaulet wholly orange red and the male genital forceps are distinctly shorter and broader. The golden pollen of the last two abdominal segments extends to the apex of each on the venter but fades out at the sides so that the very narrow hind margins of these segments above ap- pear blackish in most views. This approaches the color pattern of canadensis, but in the latter species the hind lobe of the calypter is whitish and the front pulvilli in the male are small.

Belvosia matamorosa n. sp.

A moderate-sized species allied to elusa Aldrich, which I have not seen. From the description of the latter (Proc. U.S.N.M., 73 : 25) the present species differs in lacking median marginals on the first abdominal segment and the two following ones are more extensively pollinose. Additional differences are listed below.

Male: Front at vertex 0.40 of head width, diverging immediately forward into facial angle ; parafrontals gray pollinose on an apparent blackish ground color; frontal vitta obscurely reddish, much nar- rower than parafrontal ; ocellars and orbitals absent ; outer verticals about one-half as long as inner pair ; frontals in three irregular rows, with about three bristles beneath antennal base diverging widely on upper part of parafacials, the latter also bear a few small black hairs directly below the lowermost bristles ; face, parafacial and cheek with dense satiny white pollen on yellow ground color; vibrissae about the length of second antennal segment above oral margin; facial ridge with rather stout bristles on lower three-fifths ; first two antennal segments red, third mostly black and about four times length of second ; arista brownish, moderately thickened to beyond

4 Bulletin of the Brooklyn Entomological Society XLVl

middle, thence very slender to tip; palpus stout, red, beset with numerous black hairs, haustellum short, thick, labella large and fleshy ; cheek slightly over one-third eye height, clothed with fine black hairs ; back of head flattened, cinereous, wholly pale-haired.

Thorax black, the posterior angles, scutellum and humerus red- dish brown, thinly gray pollinose above to base of scutellum, latter subshiny but showing thin tawny pollen in a flat rear view ; dorsal vittae narrow but rather well defined; sternopleurals four. Legs black ; hind tibia ciliate with one larger bristle in the row ; pulvilli subequal to length of last tarsal segment. Wing blackish basally and long costal margin to tip of first vein, thence paler to apex in- cluding the broad posterior margin ; apical cell open well before wing tip; costal spine vestigial; veins including costa blackish; sub- epaulet wholly orange red, epaulet deep reddish; calypter brown.

Abdomen black, with changeable gray pollen on second segment above, which extends thinly to the hind margin when viewed in an oblique rear angle ; third segment with somewhat denser pollen over entire surface above interrupted by a dark median stripe and con- tinuing down sides becoming heavier on venter; fourth segment covered with dense pale yellow pollen to apex; last two segments each with a marginal row of bristles ; genital forceps blackish, short and triangular as viewed from the rear, tips separated but not divergent; accessory process reddish, slightly shorter and thicker than forceps in profile, tip rounded ; fifth sternite rather small and retracted, lobes black beset with moderately coarse black hairs along the inner basal margin.

Length, 12 mm.

Holotype male, Matamoros, Pueb., Mexico, September 8, 1947 (F. A. Cowan and M. R. Wheeler).

Belvosia villaricana n. sp.

Abdominal segments one to three subshining black and without pollen except on venter, the fourth brownish in ground color, its surface above and below covered with thin changeable pale pollen from base to apex ; sternopleuron and cheek pale-haired.

Female: Front at vertex 0.33 of head width ; parafrontal yellowish gray pollinose to vertex but somewhat blackish on upper half when viewed from above ; frontal vitta red, narrowed toward vertex but at mid front nearly equal to the width of one parafrontal ; verticals two pairs, stout; ocellars absent; one reclinate and two or three proclinate orbitals ; frontals in a single row ; hairs beneath lower frontal bristles pale and delicate ; face including sides and cheeks pale yellowish white pollinose on yellow ground color ; facial ridges

Feh., 1951 Bulletin of the Brooklyn Entomological Society


with four or five bristles and some delicate white hairs outside of latter ascending to or slightly above the middle ; vibrissae nearly the length of second antennal segment above oral margin; basal an- tennal segments brownish red, the third black, about one and one- half times length of second ; arista brown, moderately flattened but slender-tipped ; palpus red, stout ; cheek barely one-third eye height ; beard pale yellowish white.

Thorax black, thinly gray pollinose above, hind angles and scutellum dark brown with tawny pollen ; five lateral scutellars on one side and four on the other, besides a short but stout preapical pair; sternopleurals four. Legs black; coxae and front femora in part pale-haired ; pulvilli hardly as long as last tarsal segment. Wing dark brown, calypter opaque blackish ; costal spine vestigial ; subepaulet wholly orange red, epaulet darker red.

Abdomen as mentioned above; two to three pairs of median marginals on first segment and four on second ; last two segments each with a marginal row ; venter mostly subshining but with some pale pollen visible along median line of the three basal segments when viewed from behind.

Length, 14 mm.

Holotype female, Villarica, Paraguay, January, 1939 (F. Schade).

Gaediophana monnula n. sp.

A moderately large black species, with the thorax thinly gray pollinose above, the wings blackish near base and the ocellar bristles uniformly absent.

Male: Front at vertex 0.32 of head width, diverging from upper fourth into facial angle ; head pollen gray, thinner on parafrontal which appears blackish, but in most views the pollen extends to the vertex ; frontal vitta deep red, much narrower than one parafrontal ; verticals two pairs; frontal row doubled, the inner or main row strongly divergent anteriorly in three or four bristles beneath an- tennal base; parafacial narrower than clypeal width, black-haired on about outer third from lowest frontals to cheek groove and with a more or less differentiated median row of stronger bristly hairs ; epistoma as wide as clypeus and bowed forward from plane of lat- ter ; vibrissae nearly on oral margin ; facial ridges strongly ciliate to upper third or more ; antenna almost as long as face, basal segments usually with a slight reddish tinge, third wholly black, about three and one-half to four times length of second ; arista black, thickened on proximal three-fourths, second segment elongate, the first short ; cheek hardly one-third eye height, clothed with fine black hairs; palpus yellow but at times darker or brownish basally ; haustellum

6 Bulletin of the Brooklyn Entomological Society ^LVI

subequal length of palpi, stout, labella fleshy; eyes thickly pilose; back of head flat, cinereous, thickly clothed with pale hairs.

Thorax black, the hind angles and scutellum brownish, latter with thin tawny pollen, subshiny in most views ; dorsal vittae narrow but distinct. Chaetotaxy : acrostichal 3,3 ; dorsocentral 3,4 ; intraalar 3 ; supraalar 3 ; presutural 2 ; notopleural 3 ; humeral 4 ; sternopleu- ral 2,2; pteropleural 1-2 (smaller than sternopleural) ; intraposta- lar differentiated ; postalar 3 ; scutellum with 3 strong lateral, 1 decussate smaller apical and 1 discal pair, besides numerous erect coarse bristly hairs on disc ; postnotal slope bare.

Legs black ; mid tibia with a row of four or five bristles on basal half of outer front side ; hind tibia irregularly ciliate on outer poste- rior side; claws and pulvilli longer than last tarsal segment.

Wing reaching well beyond apex of abdomen, subhyaline except near base ; first vein bare, third with two or three hairs near base ; apical cell open far before wing tip; hind cross vein in plane of apical cross vein and joining the fourth about one-third the dis- tance from bend to small cross vein ; costal spine vestigial ; epaulet and calypter black.

Abdomen subshining black with a more or less distinct reddish tinge in the ground color on sides, viewed in a flat rear angle the surface above is covered with thin lusterless tawny pollen; last three segments with long erect hairs that become coarser or bristle- like on the median area of second and third, each of which bears a pair of differentiated discals ; one pair of median marginals on first two segments, a marginal row on third and fourth besides a discal row on last; genital segments small; wholly black; forceps moderately elongate and slender, tips separated but not divergent, each bearing a minute basally directed spine on outer side shortly before apex, hind margin almost straight in profile ; accessory proc- ess triangular, as broad as long, polished black, with anterior mar- gin fringed with fine black hairs ; fifth sternite black, deeply excised, the lobes sparsely clothed with rather short hairs along inner mar- gin.

Female: Front at vertex 0.33 of head width; one reclinate and two proclinate orbitals ; third antennal segment about two and one- half times length of second ; abdomen short, more broadly ovate and flattened above than in male ; claws and pulvilli shorter than last tarsal segment.

Length, 9.5-12 mm.

Holotype male and allotype female, Rio Frio, Mex., Mexico, Sep- tember 3, 1947 (F. A. Cowan and M. R. Wheeler). Paratypes: 11 males and 5 females same data as type; 16 males and 11 females.

Feh., 1951 Bulletin of the Brooklyn Entomological Society


Nochixtlan, Oax, Mexico, September 6, 1947 (F. A. Cowan and M. R. Wheeler).

Siphosturmia maltana n. sp.

Similar to the genotype {rostrata Coquillett) but the abdomen is almost wholly reddish in ground color; also the male front is dis- tinctly wider and the last abdominal segment in the female is more elongated, slightly exceeding the combined length of the two pre- ceding segments.

Male: Front at vertex 0.38 of head width, about equibroad on upper third thence widening gradually into facial angle ; pollen on head wholly grayish white; frontal vitta deep red, narrower than one parafrontal; frontal rows doubled before mid front, main or inner row strongly divergent in three or four bristles beneath an- tennal base, uppermost two bristles stout and reclinate; ocellars and inner verticals strong ; basal antennal segments red, third black- ish, hardly one-third longer than second ; arista black, shorter than antenna and thickened on basal two-thirds ; clypeus hardly de- pressed, epistoma bowed forward from clypeal plane ; facial ridges flattened, with a few bristles next to vibrissae, which are near oral margin; parafacial bare below frontals, one-half clypeal width on lower extremity ; haustellum slender, tapering apically, slightly over one-half head height ; labella slender slightly elongate ; palpi yellow, bowed and but little thickened apically ; eye bare ; cheek barely one- fourth eye height.

Thorax black, with moderately dense gray pollen tinged with yel- low on mesonotum, latter with four dark vitta before suture and five behind ; acrostichal 3,3 ; dorsocentral 3,4 ; presutural 2 ; intra- alar 3 ; supraalar 3 ; intrapostalar differentiated ; sternopleural 4 ; pteropleural 1-2 (smaller than sternopleural) ; scutellum reddish, with 3 large lateral bristles besides 1 smaller decussate apical and 1 discal pair ; calypters white. Legs black ; hind tibia ciliate ; claws and pulvilli moderately elongate. Wings hyaline ; costal spine mi- nute ; first posterior cell open far before wing tip ; cubitulus obtuse angulate, without stump or fold ; epaulet black.

Abdomen red with a dark median vitta, which widens gradually towards base and is more or less obscured by rather dense whitish pollen on last three segments above; one median marginal on first two segments, a row of about 10 stouter bristles on the narrow sub- shiny hind margin of third segment and several irregular rows on shining apical half of fourth above ; genitalia small and retracted in repose ; forceps shining brown, short, rather narrow from base to tip and straight in profile, divided apically but not divergent ; acces-

8 Bulletin of the Brooklyn Entomological Society ^LVI

sory process shorter, subtriangular ; fifth sternite small, with a median V-shaped incision.

Female: Front at vertex 0.42 of head width, hardly any wider downward; two proclinate orbitals and outer verticals present; third antennal segment about one-fourth longer than second ; arista thickened on proximal three-fourths; abdomen strongly arched in profile, last segment elongate, pointed, with numerous erect short bristles above except on basal margin ; genitalia tubular, protrusile, terminating in a subchitinized blunt tip ; claws nearly as long as last tarsal segment, pulvilli distinctly shorter ; otherwise as in male.

Length, male, 10-11 mm.; female, 7.5-8 mm.

Holotype male and allotype female, Malta, Montana, July 26, 1947 (F. A. Cowan and M. R. Wheeler). Paratypes : 20 males and 6 females, same data as type.

Guerinia trudis n. sp.

Differs from the genotype (simulans Meigen) mainly in the pe- culiar structure of the male fifth abdominal sternite; the lobes of the latter are deeply excised near the middle of each inner margin and bear a dense vestiture of short coarse hairs at the base.

Male: Front at vertex 0.31 of head width; parafrontals and up- per half of parafacials pale yellow to golden pollinose, face, includ- ing lower part of parafacials and cheeks grayish white to subsilvery ; frontal vitta narrow, deep brown ; frontals descending to middle of face; facial ridges weakly bristled on lower third or less; antenna wholly black, second segment about one-half length of third ; arista black, thickened on basal half; palpus reddish; cheek nearly one- fifth eye height ; eye bare. Thorax and scutellum black, with rather dense gray pollen which usually shows a slight brassy tinge on mesonotum ; sternopleurals 3 ; acrostichals 3,3 ; dorsocentrals 3,3 ; scutellum with 3 lateral, 1 upturned strong apical and 1 smaller dis- cal pair; calypter white. Legs black, moderately long; hind tibia not ciliate ; front pulvillus subequal to combined length of two apical segments. Wings subhyaline ; first posterior cell narrowly open far before wing tip ; fourth vein with a wrinkle or fold at bend, latter subrectangular ; last section of fifth vein a little less than one-half length of preceding ; costal spine small. Abdomen long ovate, black with a distinct reddish tinge in ground color on sides and venter, segments two to four gray pollinose on basal half or more; inter- mediate segments usually without discals ; first segment with one and second generally with two median marginals ; third segment with a marginal row of about ten and fourth bristly on apical half above; genital segments rather small and retracted within tip of

Feh., 1951 Bulletin of the Brooklyn Entomological Society


abdomen ; forceps united, broadly ovate at base tapering distally to a slender, acute beak; hind surface of base concave and thickly clothed with bright yellowish hairs; fifth sternite as mentioned above, the fourth thickly beset with bristly black hairs.

Female: Front at vertex 0.35 of head width; pollen on head at times wholly gray but parafrontals usually more or less golden; two proclinate orbitals and outer verticals well developed ; second antennal segment fully two-thirds length of third, sometimes slightly reddish at apex; claws and pulvilli shorter than apical tarsal seg- ment ; otherwise, similar to male.

Length, 6-10 mm.

Holotype male and allotype female, Panguitch, Utah, June 15, 1948 (G. F. Knowlton and S. L. Wood). Paratypes: 10 males and 9 females, same data as type; 1 male, “Idaho, Aug. 10, ’19”; and 1 male. Long Valley, Alpha, Ida., June 24, 1934 (Chas. H. Martin). In the California Academy of Sciences Collection, 5 males and 2 females, Yosemite, Cal., 3880-4000 ft., June 12-17, 1931, without collector’s label, and 2 males. Hallelujah Jet., Lassen County, California, July 4, 1949 (J. W. MaeSwain).

Unusual Cockroach: During 1942, the brown banded cock- roach, Supella supellectilium (Serv.) was found to be infesting a home at Logan, Utah. The previous renter in this house had moved up from Texas, a year or so before, evidently bringing this pest along. Roaches which were sent in from a home at Payson, Utah, during August of 1943, were identified by Dr. H. K. Townes as belonging to this same species. Other specimens of this brown banded species also were called to my attention; these had been collected in Salt Lake City during 1945. Fortunately, this active species has not, as yet, become one of our “common” household pests. G. F. Knowlton, Logan, Utah.

10 Bulletin of the Brooklyn Entomological Society ^LVI


By G. B. Fairchild, Ancon, Canal Zone.

In connection with taxonomic work on Phlebotomus it was found necessary to consult a series of recent papers by Rapp, Rapp and Cooper (1944—1946) and Enderlein (1935-1937), dealing with the family Psychodidae, Rapp’s papers consist of a list of genera, with genotypes, for the world and check lists of the species of the world by zoogeographical regions. The generic list is’ compiled largely from' Enderlein (1935, 1937), with some additions. Enderlein’s final paper, the only one available to me, is a generic revision of the whole family Psychodidae, with keys and descriptions of many new genera. The following additions and corrections should be noted, though by no means all names have been carefully checked.

Phlebotomiella Meunier, 1906, Le Naturaliste (2)20 (Annee 28) : 103. Monotypic for Phlebotomus tipulijormis Meun. 1905 (Ann. Mus. Nat. Hungarici 3 : 254, PI. VI figs. 14, 15, 16). Fossil in Baltic amber. The original description and figures show an in- sect with wings reminiscent of Phlebotomus, the radius pectinately branched, but forks of R2 and R3 very close together and far distal on the wing. In terms of Phlebotometry, alpha and beta together are much less than gamma, while delta is a minus quantity. The antennae are Phlebotomus-\\ke with a long third segment. The palpi are shown as 4 segmented, the terminal segment short. I be- lieve either that the first segment was overlooked or the true ter- minal or fifth segment was lost at the time of preservation. The proboscis, although described as protruding, hardly seems suffi- ciently developed as shown in the figure to have belonged to a haematophagous insect. The male genitalia, although not very clearly described or figured, seem to have been quite similar to Phlebotomus, apparently with two terminal and a median spine on the style, terminal spines on the lateral lobes and protuberant struc- tures of some sort on the inner aspect of the coxites. Annandale (1910) mentions the genus and doubts its distinctness from Phlebotomus. The name appears to have been overlooked by sub- sequent students of the family.

Phlebotomus Rondani 1840. The spelling of this name is still a point of contention. Loew as long ago as 1845 (Dipt. Beitr. I.) said that “the name must be Latinized but not Italianized” and changed

^ Paper No. 2570 of the Scientific Journal Series, Agricultural Experiment Station, St. Paul, Minnesota.

Feh., 19S1 Bulletin of the Brooklyn Entomological Society 11

the spelling from Flehotomus to Phlehotomus. As has been noted by others (Rapp 1944, Brues 1944) Rondani himself was thoroughly inconsistent in his spelling, using Hehotomus and Phloebotomus as well as Phlehotomus and Plebotomus. His spelling of other names appears to have been equally erratic, as in 1856 (Dipt. Italicae Prodr. 1 p. 178) Psychoda phalaenoides is written Psicoda fale- noides. It is the writer’s opinion that Phlehotomus is the preferable orthography, even should it be necessary to suspend the rules to accomplish this.

A number of names have been placed as synonyms or subgenera of Phlehotomus and since there is, at present, a tendency to split the genus into an increasing number of groups, it may be well to list the available names and synonyms here.

Philaematus Loew 1845, Dipt. Beitr. 1, pp. 8-9, figs. 14, 15. Monotypic for P. pungens Loew. Fossil in Copal. From the figures this is a species of Phlehotomus. Loew’s remarks lead one to suppose that the specimens, a male and a female, are very well preserved, and that an examination of the genitalia and other struc- tures now used for classification might enable the species to be asso- ciated with some recent subgenus. It is possible that Meunier (1905, Rev. Sci. Borbonnais, 204-209) has placed pungens in Phlehotomus, but I have not seen this paper.

Cyniphes Costa 1843. Ann. Acad. Aspir. Natural. 1 : 4. With molestus Costa. The name is generally listed as a synonym of P. papatasi, but I have not seen the original publication.

Haemasson Loew 1844, Stett. Ent. Zeit., 5 : 115, with H. minutus Lw. The insect is recognizably figured, and is without doubt a synonym of P. papatasi, where it has been placed for many years.

Pophlehotomus Cockerell 1920, Ann. Mag. Nat. Hist. (9), 6, 212 fig. P. connectens Cock, sole species. Fossil in Burmese amber. Enderlein (1937) lists this as a full genus in the Tribe Mormiini on p. 98 and also, as Puphlehotomus, as a subgenus of Phlehotomus on p. 109. I do not believe it is especially closely related to Phlehotomus, and it is certainly not a subgenus. From the available descriptions and figures, it seems to stand between the Tricho- myiinae and Phlehotominae, but we know too little of its structure as yet.

Lutzomyia Franca 1924, J. Sci. Mat. Phys. Nat. Lisboa (3) 17: 10. Nom. nov. pro Lutzia Franca 1920, nec Theobald 1903. The type of Lutzia Franca was Phlehotomus longipalpis Lutz, hence the new name takes the same type, not argentipes Annan, as Rapp (1945) suggests. Lutziola Strand 1932, Lutziomyia Cordero, Vogelsang and Cossio 1928 and Pransaia Dyar and Tovar 1926,

12 Bulletin of the Brooklyn Entomological Society XLVI

were also proposed as substitute names for Lutsia Franca and hence take the same type.

Shannonomyina Pratt 1947, Proc. Ent. Soc. Washington 49: 86. Nom. nov. pro Shannonomyia Dyar 1929 (July), nec Shannono- myza Alexander 1929 (January) (Tipulidae). Phlebotomus

panamensis Shannon.

Sergentomyia Franca and Parrot 1920. The name was proposed to replace N ewsteadia Franca 1919 (nec Newsteadia Green 1902), which was proposed to include six mediterranean species considered subgenerically distinct from P. papatasi. Later in 1920, Franca designated P. minutus Rond, as genotype of Sergentomyia. Rapp’s designation of P. papatasi as genotype of N ewsteadia is quite un- warranted. Prophlehotomus Franca and Parrot 1921, was split off from Sergentomyia, without type designation, but including P. minutns, the. previously designated genotype of Sergentomyia, so that the name falls as a synonym of Sergentomyia. Rapp’s action in selecting P. pertiirbans de Meijere 1909 seems unnecessary and would probably not alter the synonymy as Theodor (1948) lists perturbans as an unrecognizable species, probably belonging in Sergentomyia. According to Theodor (1948), Neophlebotomus Franca and Parrot 1920, (Type P. malabaricus Annand. 1910) cannot be separated from Sergentomyia, while Brumptius Nitzu- lescu 1931, is also a synonym, being isogenotypic.

The following names were proposed as subgenera with the types indicated by original designation.

Brumptomyia Franca and Parrot 1921. Type P. brumpti Larr. 1920.

Sintonius Nitzulescu 1931. Type P. hospitii Sinton 1924.

Larroussius Nitzulescu 1931. Type P. major Annand. 1910.

Adlerius Nitzulescu 1931. Type P. chinensis Newst. 1916.

Pintomyia Costa Lima 1932. Type P. fischeri Pinto 1926.

Evandromyia Mangabeira 1941. P. infraspinosus Mang. 1941.

Psychodopygus Mang. 1941. Type P. unisetosus Mang. 1941.

Viannamyia Mang. 1941. Type P. tubercidatus Mang. 1941.

Pressatia Mang. 1942. Type P. triacanthus Mang. 1942.

Castromyia Mang. 1942. Type P. castroi Barr, and Gout. 1941.

Dampfomyia Addis 1945. Type P. anthophorus Addis 1945.

Paraphlebotomus Theodor 1948. Type P. sergenti Parrot 1917.

Synphlebotomus Theodor 1948. Type P. martini Parrot 1936.

Euphlebotomus Theodor 1948. Type P. argentipes Ann. and Brun 1908.

Anaphlebotomus Theodor 1948. Type P. stantoni Newst. 1917.

Fel., 1951 Bulletin of the Brooklyn Entomological Society 13

Australophlebotomus Theodor 1948. Type P. brevi fills Tonn. 1935.

Spelaeophlebotomus Theo. 1948. Type P. qiqas Parrot and Schwetz 1937.

Spelaeomyia Theo. 1948. Type P. mirabilis Parrot and Wanson 1939.

The paper by Theodor (1948) was, of course, not available to Rapp when he compiled his list, but Costa Lima’s 1932 paper should have been consulted for the earlier names, while Mangabeira’s pub- lications all appeared several years before Rapp’s list.

Tinearia Schellenberg 1803. This name is listed by both Ender- lein and Rapp as valid with Trichoptera fuliginosa Meigen 1804 as genotype and Ulomyia Haliday in Walker 1856 as a synonym. Coquillet, however, (1910) designated Psychoda alternata Say 1824 as genotype, and placed Tinearia as a synonym of Psychoda. An examination of Schellenberg’s original publication (Genres des Mouches Dipteres, representes en XLII Planches projettees et dessinees par Mr. J. R. Schellenberg et expliquees par deux ama- teurs de I’Entomologie. Zurich (1803) indicates quite clearly that Schellenberg is responsible only for the plates and the names appear- ing on them. Apparently the publishers felt some sort of text necessary for they appear to have secured the services of two anonymous “Amateurs” to supply this deficiency. There seems to have been no contact between Schellenberg and the authors of the text. The authors of the text state in their introduction that they intend to follow the classification of the “immortal Fabricius.” They say also that the plates were already printed before they were asked to supply the text, and hence no changes could be made in the former. The result is that the plates often bear no names, generic names only, or names different from those given in the text. In the case of the Psychodidae the plate XL bears the supposed generic name Tinearia, but no specific name. The figures are numbered lA, a, b, c, 2B and d. The explanation of the plate is on page 23 and gives the name as Tipula Phalaenoides Fabr. Ent. Syst. IV p. 251 No. 85. This covers figures lA, a, b, and c of the plate. Figures 2B and d are named as Tipula hirta Fabr. Ent. Syst. IV p. 251 no. 84. The plates are very poor. No mention of the name Tinearia appears in the text.

Since the text of this work was not by Schellenberg and the name Tinearia is nowhere associated with a specific name either by Schellenberg or the anonymous authors of the text, it seems that the name must be considered a genus without species. Coquillet’s

14 Bulletin of the Brooklyn Entomological Society XLVI

designation of alternata Say thus is valid, since the naming of the figures in the plate without using the generic name on the plate does not restrict Tinearia in any way. Enderlein seems to have been unaware of the dual authorship of this work, or of the previous designation of a genotype by Coquillet. Rapp, in following Ender- lein, seems to have been unaware that Ulomyia Hal. (not Walker) 1856 was a substitute name for Saccopteryx Hal. 1839 preoccupied. Both names are listed separately as synonyms of Tinearia by Ender- lein and Rapp on the basis of isogenotypy. Ulomyia is considered by recent workers (Coe 1945) as a subgenus of Pericoma.

Panimerus Eaton 1913 Trans. Linn. Soc. London (2) 15: 425- 427. Type by original designation “Panimerus hirtus (Linn) (=notabilis Eaton)”. Enderlein (1937) and Rapp give P. scotti Eaton 1913 as type, presumably because Enderlein (1935) desig- nated notabilis Eaton as genotype of Lepiseoda End. 1935, a genus which he sank (1937) under Panimerus with the statement that Tonnoir considered scotti Eaton 1913 and notabilis Eaton 1893 synonymous.

Lepidopsychoda Edwards 1928, Insects Samoa, Pt. VI, fasc. 2, pp. 7.1-72, fig. 10, Type by original designation L. tineiformis Ed- wards 1928. Also includes Brunettia trimicra Edw. This genus is not mentioned by either Enderlein or Rapp.

Mesopsychoda Brauer, Redtenbacher and Ganglbauer 1889 (Akad nauk, SSSR Leningrad, Memoires, Ser. 7, Vol. 36, No. 15, pp. 1-22, 2 plates) with dasyptera sole species. Fossil. Jurassic. East Siberia. I have not seen the original description but Hand- lirsch (1908. Die fossilen Insekten, p. 629, pi. 51, fig. 4) has re- described and figured the specimen. It is small, 3.6 mm. long, with the hairy wings folded tent-like over the abdomen. What can be made out of the venation is not very like modern Psychodids, but the material is probably not adequate for detailed comparisons. Handlirsch also creates the provisional genus Psychodites for two species, kenngotti Giebel and egertoni Brodie which he believes may be Psychodids, though from his figures there is little to sup- port this view except that the wings are more or less hairy.

Parabrunettia Brunetti 1911. According to Tonnoir (1939) the history and synonymy of this name is as follows : Parabrunettia has as type P. squamipennis Brun., designated by Brunetti in 1912. But this species is a true Brunettia, hence Parabrunettia = Brunet- tia. Parabrunettia of Enderlein, with B. indica Eaton 1913 as type is subgenerically distinct, and Tonnoir proposes for it the name Trichobrunettia. He also points out that not only B. indica Eaton but also Parabrunettia 9-notata Brun. and Psychoda duripuncta

Fel., 1951 Bulletin of the Brooklyn Entomological Society 15

Curran are synonyms of Brunettia alhonotata Brun. 1908, a more or less tropicopolitan species. Rapp apparently did not see this paper, as neither the generic nor specific synonymies nor the name Trichobrunettia appear in his lists.

Posthon Loew 1845 (Dipt. Beitr. 1 : 9-10) Type gracilis Loew. 1850.

Phalaenomyia Loew 1845 (l.c.) no species named.

These two names, based on fossil material in amber, were placed respectively as synonyms of Sy corax and Trichomyia by Haliday, in Walker 1856, where they have remained. At least the first is nomenclatorially available, and should the type ever turn up, might well preoccupy later names. Their descriptions are meagre, con- sisting of brief comparisons with Diplonema Loew. Giebel (1856) has discussed all the fossil Psychodidae named at that time, while Meunier (1905) has described and figured a considerable number of amber forms and given keys to the known fossil genera and spe- cies.

Phalaenula Meigen 1800. Coquillet (1910) designated Tri- choptera ocellaris Meigen 1804 as genotype. Eaton in 1904 had erected the genus Clytocerus, without type designation, but includ- ing by citation Pericoma ocellaris (Meigen) and P. dalei Eaton and a figure of the former. Enderlein in 1935 designated P. dalei Eaton as genotype of Clytocerus, but in 1937 he gives ocellaris Meigen as genotype, in which he is followed by Rapp. Stone (1941) considers Clytocerus to be a synonym of Phalaenula, ap- parently the only correct procedure under the rules if Meigen’s 1800 names are accepted, while Coe 1945, recognized Clytocerus as valid with ocellaris and dalei.

Termitadelphos Holmgren 1905 Type silvestrii Holmg.

Termitodipteron Holmgren 1905 Type Wasmanni Holmg.

These two genera are not mentioned by Rapp or Enderlein, though Tonnoir (1929 p. 2 footnote) considers the first as a pos- sible synonym of Psychoda, the second provisionally in the Tricho- myiinae.

Eatonisca Meunier 1905. Type E. tertiaria Meun. 1905. Eossil in Baltic amber. This genus is placed next to Horaiella Tonn. by Enderlein, but omitted by Rapp, although he includes other fossil genera. I have seen the original description.

Eutonnoiria Alexander 1940 (Rev. Ent. 11: 794) Type Bru- chomyia edwardsi Tonn. 1939. This name seems to have been overlooked by Rapp.

“Diplomia Annandale” Rapp and Cooper 1945 p. 211. This is a misspelling of Diplonema Annand. 1908, which name is preoc-

16 Bulletin of the Brooklyn Entomological Society XLVI

cupied by Diplonema Loew 1845, as discovered by Annandale two years later when he renamed the genus Briinettia. Rapp and Cooper list the single species sup erst es under Diplomia rather than under Brunettia, whose genotype it is. Diplonema Loew with type buceras is correctly listed by both Rapp and Enderlein.

There are a distressingly large number of minor errors and omis- sions in the series of papers by Rapp, but there seems no point in going into details. Dates are often omitted and references are in too many cases quite inaccurate. For example, in Rapp and Cooper 1945, pp. 214-215, N emo palpus australiensis Alexander was de- scribed in Proc. Linn. Soc. N.S. Wales 53: 293-294, 1928, and not in the Federated Malay States Museum Journal 14: 65, as they have it. N. orientalis Fdwards 1928, which was described in the Federated Malay States Museum Journal 14: 65, is not listed by Rapp. In the same genus N. tertiariae (Meunier 1905) and N. molophilinus (Fdwards 1921), two fossil species, are nowhere mentioned by Rapp, while N. zelandiae Alex. 1921 is misspelled zelandicus.

In going over the faunal lists, no attempt was made at a com- plete check, but the following omissions should be noted. The genus Maruina is not listed as North American by Rapp, in spite of dehnite statements by Tonnoir (1929, 1934) and Fdwards (1929) that Pericoma californiensis Kellogg represents the early stages of Maruina lanceolata Kinc. and that Trichomyia unipunc- tata URseun. = Maruina, and by Johannsen (1938), who lists and gives a key to six species, three of them from the United States. Rapp places two of these latter in Trichomyia and omits the third entirely. True Trichomyia occurs in the United States also, as I have examined specimens of T, urbica Curtis in the U.S.N.M. from Virginia. I have also compared specimens of Maruina lance- olata Kinc. with the original descriptions of Maruina by F. Muller and consider them congeneric. It may be well to note here that the figure of the wing of Maruina in Curran’s North American Diptera, 1934, p. 79 is a species of Sy corax.

The listings of the species of PhlebotomusxiVQ very far from com- plete. About a dozen species described previous to Rapp’s paper on the African Psychodidae are missing, while P. troglodytes Lutz 1922, a Brazilian species, is listed as P. troglodytes Nitzulescu 1930 from Tunisia. The paper quoted is actually a discussion of differ- ences between troglodytes Lutz and brumpti Larr. The Oriental lists are about equally incomplete and there are minor slips, such as the crediting of P. barraudi S inton 1929, to Yao and Wu and P. nicnic Banks